Cyperaceae. This family usually has 3-angled stems, which is most obvious immediately below the inflorescence. Some Cyperaceae species have cylindrical stems similar to grasses. In this case, the spikelets differ from Poaceae in having flowers mostly helically arranged (grasses have distichous spikelets) and the flowers differ in lacking glumes and paleas. The perianth may be present in the form of barbed bristles, which is unique to Cyperaceae. Characteristics of the common genera are reported below.

1. Eleocharis palustris. Common spikerush. Rhizomatous perennials often mat-forming if not bunched, distinguished from other spikerushes by the combination of its lenticular achenes that bear 2 stigma branches, stems over 15 cm tall, and perennial rhizomatous habit. Eleocharis is distinguished from other Cyperaceae genera by inflorescences of a single terminal spikelet and achene with a persistent pyramidal style.

2. Scirpus microcarpus. Panicled bulrush. A rhizomatous perennial often forming dense stands along with other wetlands species. This species is distinguished from other bulrushes by its small spikelets in an open panicle subtended by several leafy bracts, by rusty or reddish lower leaf sheaths, and by having a lenticular (not 3-angled) achene. Scirpus is very closely related to Eleocharis and is distinguished by having more than one spikelet terminate a flowering stem and flower with an ovary having a thin style.

3. Scirpus validus (Schoenoplectus tabernaemontani). Softstem bulrush. A rhizomatous perennial forming extensive and dense stands in ponds and along ditches. Although with cylindrical stems, note that the inflorescences, spikelets, and flowers are similar to Scirpus microcarpus.

4. Cyperus esculentus. Yellow nutgrass. A rhizomatous perennial and often an aggressive colonizer. Cyperus is the only genus of Cyperaceae in Montana that commonly could be confused with the Poaceae because of its laterally flattened distichous spikelets. The spikelets occur in heads surrounded by radially arranged leaves, something never seen in grasses. The 3-angled stem is indicative of Cyperaceae and not Poaceae. Although the spikelet is very grass-like, glumes and paleas are lacking.

5. Carex filifolia, threadleaf sedge. Bunched and mat-forming perennials of typically open dry sites such as in the sagebrush steppe, the inflorescence is a sessile terminal spikelet, perigynia surround an elliptical achene. Staminate flowers terminate the distal end of the spikelet whereas the pistillate flowers are borne towards the base. Carex with an inflorescence comprising a single spikelet are usually dry site inhabiting. Carex is distinguished by monoecious flowers, the pistillate of which produce a perigynium.

6. Carex hoodii, hood sedge. Bunched perennials typically of mountain meadow settings, the inflorescence comprises more than several sessile and tightly clustered spikelets, the winged perigynia surround an elliptical achene. Each spike terminates with staminate flowers (note the thin whitish filaments if anthers are not present) whereas the pistillate flowers are borne below the tip of each spikelet to its base. Carex with an inflorescence comprising sessile congested spikelets are usually dry site to dry meadow inhabiting.

7. Carex paysonis, payson sedge. Bunched to single-stemmed perennials of subalpine to alpine meadows, the inflorescence comprises several or more pedicellate spikelets where the distal-most spikelets bear only staminate flowers and the basal-most only pistillate flowers. The perigynia surround 3-angled (trigonous) achenes. The broad dark flower bracts are distinctive of this species and often render the entire spikelet distinctively black or dark brown in color. Carex with an inflorescence comprising pedicellate spikelets that are well spaced are usually wet meadow to wet site inhabiting.

Juncaceae. The small flowers of Juncaceae have parts in 3’s, unlike other graminoid families. The fruit is also different from other graminoid families in being a capsule that splits open along three sutures and spills out three to many seeds. The seed is the dispersal unit.

8. Juncus balticus. Baltic rush. A strongly rhizomatous perennial with stems sometime approaching a meter tall, a colonizer of disturbed open grassy sites at mid elevations. Juncus produces leaf sheaths with our without well developed leaf blades but in any case the leaf sheaths are open and the collar, throat, and margins of the leaf blades are glabrous.

9. Juncus ensifolius. Swordleaf rush. A rhizomatous perennial common in seeps, along roadsides, etc., the laterally flattened leaves (folded flat along the midrib in fresh condition) and styles that protrude distinctly beyond the tepals are characteristic of this species.

10. Juncus bufonius. Toad rush. A bunched annual with stems typically less than 25 cm tall, of moist meadows, the solitary flowers scattered along the inflorescence rachis is distinctive of this species. Most Juncus have flowers congested into heads or at least near the distal branch ends.

11. Luzula parviflora. Small-flowered woodrush. A single-stemmed perennial of moist mountain meadows. This species is one of the more common woodrushes and is distinguished from the others by its open paniculate inflorescence, small flowers, and fairly ample leaves. Woodrushes differ from rushes in having a fused leaf sheath, long hairs along the margins of young leaf blades, and only one seed per ovary compartment or carpel.

Typhaceae. The inflorescence is a terminal cylindrical spike with pistillate flowers borne below the staminate ones. The pistillate flower has a perianth of long thin hairs which serve to disperse the flower and seed.

12. Typha latifolia. Broadleaf cattail. A tall native perennial forming extensive stands, flowers are monoecious, the long-stipitate ovary is ultimately dispersed by the pappus-like perianth. The long styles flare distally into a brownish end. These style ends from both sterile and fertile pistillate flowers when packed together render the brownish coloration of the pistillate portion of the spike. The staminate flowers comprise 2-5 stamens borne from a hair-like pedicel. Typha latifolia is distinguished from other cattails in this region by its inflorescence in which the staminate portion is contiguous with the pistillate portion.

syllabus